Polystichum lepidocaulon (Scale-stem Cyrtomidictyum, 鞭葉耳蕨)

Scale-stem Cyrtomidictyum is in the genus Polystichumnative and family Dryopteridaceae , native to China, Korea, Japan and Taiwan. It is terrestrial with maximum height about 28 to 48 cm and the fronds are tufted. Its rhizome dense quilt dark brown scales, so it has the English name 'Scale-stem' Cyrtomidictyum.

Scale-stem Cyrtomidictyum is a kind of 'walking fern'. The rachises of its some fronds are prolonged and grow adventitious buds at the tip. When the buds grow bigger and bigger, the rachises droop to the ground and the buds root downwards and become young plants independently. Its Chinese name '鞭'葉 means 'whip' leaf because its prolonged rachises look like whips and that is the characteristics of Scale-stem Cyrtomidictyum.
Scale-stem Cyrtomidictyum
Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

About Scale-stem Cyrtomidictyum
Classification: Genus Polystichum; Family Dryopteridaceae 
Scientific name: Polystichum lepidocaulon (Hook.) J. Sm.
English Name: Scale-stem Cyrtomidictyum
Chinese name: 鞭葉耳蕨、鞭葉蕨
Origin: China, Korea, Japan, Taiwan
Growth environment:On the soil slope under the middle altitude forest (in Taiwan)
Growth habit: Terrestrial
Maximum height: About 28 to 48 cm
Rhizome: Short, ascending, dense quilt scales which are dark brown, ovate; frond tufted
Frond
  Stipe 13-35 cm long,  dense quilt scales which are dark brown or blackish, ovate to lanceolate
  Blade
→Pinnate, lanceolate
→25-30 cm long, 5-7 cm broad
→Groove on upper surface of the rachis; brown scales on lower surface of the rachis
→Rachises of some fronds prolonged and grow adventitious buds at tip
  Pinna
→Soft coriaceous, falcate lanceolate, acuminate at apex; entire; with auricle on the upper side of the base
→3-6 cm long, 0.7-2 cm broad; without or with very short pinna stalk
→Glabrous on upper surface; small scales on under surface
  Veins free and branched
Sori & Indusia
  Sori dorsal on veinlets, arranged in 1-3 rows on each side of costa
  Indusia very small, rounded, peltate, caducous

Scale-stem Cyrtomidictyum is native to China, Korea, Japan and Taiwan. It is terrestrial with maximum height about 28-48 cm and the fronds are tufted.
Scale-stem Cyrtomidictyum
Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

Rhizome is short, ascending and dense quilt scales which are dark brown and ovate.
Scale-stem Cyrtomidictyum
Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

Stipes are about 13-35 cm long,  dense quilt scales which are dark brown or blackish, ovate to lanceolate.
Stipe of Scale-stem Cyrtomidictyum
Stipe of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

 The blade is pinnate.
Blade of Scale-stem Cyrtomidictyum
Blade of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

The blade is lanceolate.
Blade of Scale-stem Cyrtomidictyum
Blade of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

The blade is about 25-30 cm long and about 5-7 cm broad.
Blade of Scale-stem Cyrtomidictyum
Blade of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

There is groove on upper surface of the rachis and brown scales on lower surface of the rachis.
Rachis of Scale-stem Cyrtomidictyum (upper surface)
Rachis of Scale-stem Cyrtomidictyum (upper surface)
(2022.3.29 Taiwan)

The rachises of some fronds are prolonged and grow adventitious buds at the tip.
Frond of Scale-stem Cyrtomidictyum
Frond of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

The adventitious buds can grow to be new plants.
Adventitious bud of Scale-stem Cyrtomidictyum
Adventitious bud of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

Adventitious bud of Scale-stem Cyrtomidictyum
Adventitious bud of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

Adventitious bud of Scale-stem Cyrtomidictyum
Adventitious bud of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

The pinnae are soft coriaceous, falcate lanceolate and acuminate at apex. They are entire and with auricle on the upper side of the base.
The pinnae are about 3-6 cm long and 0.7-2 cm broad and without or with very short pinna stalk.
Pinnae of Scale-stem Cyrtomidictyum
Pinnae of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

It is glabrous on upper surface and with small scales on lower surface of the pinna.
Pinnae of Scale-stem Cyrtomidictyum (lower surface)
Pinnae of Scale-stem Cyrtomidictyum (lower surface)
(2022.3.29 Taiwan)

Veins are free and branched.
Veins of Scale-stem Cyrtomidictyum
Veins of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

Sori are dorsal on veinlets, arranged in 1-3 rows on each side of costa.
Sori of Scale-stem Cyrtomidictyum
Sori of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

Indusia are very small, rounded, peltate and caducous.
Sori of Scale-stem Cyrtomidictyum
Sori of Scale-stem Cyrtomidictyum
(2022.3.29 Taiwan)

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